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Additional resources for Biochemistry. The Chemical Reactions of Living Cells. Vol.2
2 µM, is too low to measure. In contrast to animals, the resurrection plant Craterostigma plantaginenm accumulates large amounts of a 2-oxo-octulose. This plant is one of a small group of angiosperms that can withstand severe dehydration and are able to rehydrate and resume normal metabolism within a few hours. During desiccation much of the octulose is converted into sucrose. 100 However, it is not clear how the C3 unit used in the nonoxidative pathway (Fig. 17-8B) is formed if glycolysis is blocked.
Why does the glycolysis sequence begin with phosphorylation of glucose by ATP? The phospho groups probably provide convenient handles and doubtless assist in substrate recognition. There may be a kinetic advantage but also a danger. 139a Trypanosomes utilize a different type of control. The enzymes that convert glucose into 3-phosphoglycerate are present in membrane-bounded organelles called glycosomes. 139b Since inorganic phosphate is essential for ATP formation, if the Pi concentration falls too low the rate of fermentation by yeast juice is greatly decreased, an observation made by Harden and Young139c in 1906.
8 kcal mol – 1) (17-19) Glucose → 2 CO2 + 2 ethanol ∆G° = – 235 kJ mol – 1 (17-20) The Gibbs energy changes are negative and of sufficient magnitude that the reactions will unquestionably go to completion. However, the synthesis of two molecules of ATP from inorganic phosphate and ADP, a reaction 2/14/03, 11:16 AM F. Fermentation: “Life without Oxygen” (Eq. 17-21) for which ∆G’ is substantially positive, is coupled to the fermentation. 0 kJ to the values of ∆G’ for Eqs. 17-19 and 17-20. When this is done we see that the net Gibbs energy changes are still highly negative, that the reactions will proceed to completion, and that these fermentations can serve as an usable energy source for organisms.